/* Original Kojetin Laboratory design. No theme or UI framework. */
:root {
  --paper: #f7f7f2;
  --ink: #253131;
  --muted: #606c6b;
  --line: #d7ddd7;
  --teal: #277b77;
  --gold: #9c7445;
  --violet: #777e9c;
  --max: 1320px;
}
* {
  box-sizing: border-box;
}
html {
  scroll-behavior: smooth;
  scroll-padding-top: 110px;
}
body {
  margin: 0;
  background: var(--paper);
  color: var(--ink);
  font-family: Arial, Helvetica, sans-serif;
  font-size: 17px;
  line-height: 1.6;
  -webkit-font-smoothing: antialiased;
}
a {
  color: inherit;
  text-decoration: none;
}
a:hover {
  color: var(--teal);
}
button,
input,
select {
  font: inherit;
  color: inherit;
}
button,
a,
input,
select {
  touch-action: manipulation;
}
button {
  cursor: pointer;
}
button:focus-visible,
a:focus-visible,
input:focus-visible,
select:focus-visible {
  outline: 2px solid var(--teal);
  outline-offset: 5px;
}
img {
  max-width: 100%;
  height: auto;
}
h1,
h2,
h3,
p,
figure {
  margin: 0;
}
h1,
h2,
h3 {
  font-weight: 400;
  line-height: 1.13;
  letter-spacing: -0.04em;
}
p {
  margin: 0 0 1em;
}
h2 {
  font-size: clamp(32px, 3.5vw, 49px);
}
h3 {
  font-size: 24px;
}
button {
  border-radius: 0;
}
small,
.small {
  font-size: 14px;
}
::selection {
  background: var(--selection);
}
.wrap {
  width: min(calc(100% - 112px), var(--max));
  margin-inline: auto;
}
[hidden] {
  display: none !important;
}
.skip-link {
  position: fixed;
  left: 20px;
  top: -90px;
  padding: 12px 20px;
  background: var(--surface);
  z-index: 100;
}
.skip-link:focus {
  top: 12px;
}
.eyebrow {
  font-size: 11px;
  line-height: 1.6;
  letter-spacing: 0.14em;
  text-transform: uppercase;
  font-weight: 600;
  color: var(--muted);
  margin-bottom: 24px;
}
.text-link {
  display: inline-flex;
  gap: 32px;
  align-items: center;
  font-size: 14px;
  border-bottom: 1px solid var(--line);
  padding-bottom: 8px;
}
.text-link span {
  font-size: 20px;
}
.large-copy {
  font-size: clamp(20px, 2.2vw, 28px);
  line-height: 1.5;
  letter-spacing: -0.025em;
}
.site-header {
  position: relative;
  z-index: 20;
  display: flex;
  justify-content: space-between;
  align-items: center;
  max-width: 1550px;
  margin: auto;
  padding: 30px 56px 24px;
  border-bottom: 1px solid var(--line);
  min-height: 102px;
  background: var(--paper);
}
.wordmark {
  display: flex;
  align-items: baseline;
  gap: 8px;
  font-size: 30px;
  font-weight: 750;
  flex-shrink: 0;
  letter-spacing: -0.8px;
  line-height: 1;
}
.wordmark span {
  font-size: 16px;
  font-weight: 400;
  letter-spacing: -0.5px;
}
.wordmark i {
  width: 6px;
  height: 6px;
  border-radius: 50%;
  background: var(--gold);
  align-self: flex-end;
  margin: 0 0 3px 0;
}
.site-header nav {
  display: flex;
  align-items: center;
  gap: 26px;
  font-size: 13px;
}
.site-header nav a {
  position: relative;
  padding: 8px 0;
}
.site-header nav a[aria-current] {
  color: var(--teal);
}
.site-header nav a[aria-current]:after {
  content: "";
  position: absolute;
  height: 2px;
  background: var(--teal);
  bottom: 0;
  left: 0;
  right: 0;
}
.menu-toggle {
  background: none;
  border: 0;
  padding: 9px 0;
  font-size: 14px;
}
.site-footer {
  padding: 56px 0 40px;
  background: var(--paper);
  color: var(--muted);
  border-top: 1px solid var(--line);
  font-size: 12px;
}
.footer-brand {
  display: block;
  width: min(100%, 560px);
  margin-bottom: 44px;
}
.footer-brand img {
  display: block;
  width: 100%;
  height: auto;
}
.footer-grid {
  display: grid;
  grid-template-columns: 1.3fr 1.4fr 1fr;
  gap: 50px;
}
.footer-name {
  font-size: 17px;
  color: var(--ink);
  letter-spacing: -0.5px;
}
.site-footer a:hover {
  color: var(--teal);
}
.site-footer a:focus-visible {
  outline-color: var(--teal);
}
.site-footer p {
  margin-top: 12px;
}
.footer-grid > div:last-child {
  text-align: right;
}
.site-footer .footer-source {
  margin-top: 36px;
  padding-top: 20px;
  border-top: 1px solid var(--line);
}
.footer-source a {
  text-decoration: underline;
  text-underline-offset: 3px;
}
.molecular-story {
  position: relative;
}
.story-stage {
  position: relative;
  min-height: calc(100svh - 102px);
  display: grid;
  grid-template-columns: 45% 55%;
  align-items: center;
}
.hero-copy {
  padding: 56px 0 82px;
  position: relative;
  z-index: 2;
  pointer-events: none;
}
.hero-copy a {
  pointer-events: auto;
}
.hero-copy h1 {
  font-size: clamp(70px, 8.6vw, 132px);
  line-height: 0.97;
  letter-spacing: -0.065em;
}
.hero-copy h1 span {
  color: var(--teal);
}
.hero-intro {
  font-size: 18px;
  line-height: 1.55;
  max-width: 335px;
  color: var(--muted);
  margin: 30px 0 32px;
}
.molecular-figure {
  position: relative;
  align-self: center;
  width: 100%;
  min-width: 0;
}
.molecule-view {
  position: relative;
  width: 100%;
  aspect-ratio: 1;
  overflow: hidden;
}
.molecule-layer {
  position: absolute;
  inset: 0;
  z-index: 2;
  transform-origin: center;
}
.cell-scene {
  position: absolute;
  inset: 0;
  z-index: 1;
  opacity: 0;
  pointer-events: none;
}
.cell-scene .cell-artboard {
  position: absolute;
  left: 50%;
  top: 50%;
  transform: translate(-50%, -50%);
}
.cell-artboard,
.cell-art {
  position: relative;
  width: 100%;
  aspect-ratio: 1;
}
.cell-illustration {
  display: block;
  width: 100%;
  height: 100%;
  object-fit: contain;
}
.cell-labels {
  opacity: var(--cell-label-opacity, 1);
}
.cell-label {
  position: absolute;
  padding: 3px 7px;
  border-radius: 4px;
  background: color-mix(in srgb, var(--paper) 90%, transparent);
  color: var(--ink);
  font-size: var(--text-caption);
  line-height: 1.3;
}
.cell-label-nucleus {
  left: 7%;
  top: 50%;
}
.cell-label-lipid {
  left: 52%;
  top: 47%;
}
.cell-label-cytoplasm {
  left: 34%;
  top: 83%;
}
.cell-static {
  margin: 24px 0 0;
  width: 100%;
  max-width: 500px;
}
.cell-static figcaption {
  font-size: var(--text-caption);
  line-height: 1.5;
  color: var(--muted);
}
.cell-transition-enabled .cell-static,
.pdb-link[hidden] {
  display: none;
}
.molecule-view canvas,
.molecule-poster {
  position: absolute;
  inset: 0;
  width: 100%;
  height: 100%;
  object-fit: contain;
}
.molecule-view canvas {
  opacity: 0;
}
.molecule-ready .molecule-view canvas {
  opacity: 1;
}
.molecule-ready .molecule-poster {
  opacity: 0;
}
.molecule-label {
  position: absolute;
  z-index: 3;
  padding-left: 18px;
  font-size: 12px;
  line-height: 1.4;
  pointer-events: none;
  background: color-mix(in srgb, var(--paper) 85%, transparent);
  border-radius: 3px;
  padding-right: 5px;
  opacity: var(--molecule-label-opacity, 1);
}
.molecule-label:before {
  content: "";
  position: absolute;
  left: 4px;
  top: 5px;
  width: 5px;
  height: 5px;
  border-radius: 50%;
  background: var(--violet);
}
.molecule-label small {
  display: block;
  font-size: 9px;
  color: var(--muted);
}
.label-peptide {
  top: 18%;
  right: 2%;
}
.label-peptide:before {
  background: var(--teal);
}
.label-ligand {
  bottom: 24%;
  left: 1%;
}
.label-ligand:before {
  background: var(--gold);
}
.label-receptor {
  right: 8%;
  bottom: 12%;
}
.molecular-figure figcaption {
  display: flex;
  justify-content: space-between;
  flex-wrap: wrap;
  gap: 8px;
  font-size: 10px;
  color: var(--muted);
  border-top: 1px solid var(--line);
  padding: 13px 0;
}
.figure-index {
  font-size: 9px;
  letter-spacing: 0.09em;
}
.figure-note {
  flex-basis: 100%;
  font-size: 10px;
}
.motion-toggle {
  position: absolute;
  right: 0;
  bottom: -30px;
  background: none;
  border: 0;
  font-size: 10px;
  color: var(--muted);
  padding: 3px 0;
  border-bottom: 1px solid var(--line);
}
.story-steps {
  position: relative;
}
.story-step {
  max-width: 380px;
  padding: 80px 0;
}
.story-step h2 {
  font-size: clamp(38px, 4.2vw, 60px);
  margin-bottom: 25px;
}
.story-step p {
  font-size: 17px;
  color: var(--muted);
  line-height: 1.7;
}
.story-step .text-link {
  margin-top: 20px;
}
.motion-enabled .molecular-story {
  padding-bottom: 50px;
}
.motion-enabled .story-stage {
  position: sticky;
  top: 15px;
  height: calc(100svh - 30px);
  min-height: 650px;
}
.motion-enabled .story-steps {
  pointer-events: none;
  margin-top: 0;
}
.motion-enabled .story-step {
  min-height: 70vh;
  display: flex;
  flex-direction: column;
  justify-content: center;
  pointer-events: auto;
}
.motion-enabled .hero-copy {
  opacity: var(--intro-opacity, 1);
}
.motion-enabled .story-stage > .molecular-figure {
  grid-column: 2;
  grid-row: 1;
}
.motion-enabled .hero-copy {
  grid-column: 1;
  grid-row: 1;
}
.mission-section {
  border-top: 1px solid var(--line);
  padding: 105px 0;
  display: grid;
  grid-template-columns: 1fr 2fr;
  gap: 60px;
}
.mission-section h2 {
  font-size: clamp(32px, 3.6vw, 50px);
  margin-bottom: 30px;
}
.mission-section .large-copy {
  color: var(--muted);
  font-size: 23px;
  max-width: 800px;
}
.section-heading {
  display: flex;
  justify-content: space-between;
  align-items: baseline;
  gap: 30px;
  margin-bottom: 36px;
}
.section-heading h2 {
  font-size: 38px;
}
.research-preview {
  padding-bottom: 110px;
}
.research-row {
  display: grid;
  grid-template-columns: 1.1fr 1fr 24px;
  gap: 30px;
  align-items: start;
  border-top: 1px solid var(--line);
  padding: 34px 0;
}
.research-row:last-child {
  border-bottom: 1px solid var(--line);
}
.research-row h2,
.research-row h3 {
  font-size: 27px;
  letter-spacing: -0.035em;
  line-height: 1.25;
}
.research-row p {
  font-size: 15px;
  color: var(--muted);
  max-width: 420px;
  margin: 0;
}
.selected-publications {
  padding-bottom: 105px;
}
.publication-group + .publication-group {
  margin-top: 64px;
}
.publication-group > h2 {
  margin-bottom: 12px;
}
.publication-group-note {
  color: var(--muted);
  margin: 0 0 24px;
}
.publication-abstract {
  margin-top: 36px;
}
.publication-row {
  display: grid;
  grid-template-columns: 250px 1fr;
  gap: 42px;
  border-top: 1px solid var(--line);
  padding: 34px 0;
}
.publication-row:last-child {
  border-bottom: 1px solid var(--line);
}
.publication-image {
  display: flex;
  align-items: center;
  justify-content: center;
  min-height: 160px;
  background: var(--surface);
  padding: 12px;
}
.publication-image img {
  width: 100%;
  max-height: 220px;
  object-fit: contain;
}
.publication-year-art {
  background: var(--tint);
  min-height: 150px;
  display: flex;
  flex-direction: column;
  align-items: center;
  justify-content: center;
  color: var(--muted);
  font-size: 52px;
  font-weight: 400;
  letter-spacing: -2px;
}
.publication-year-art span {
  font-size: 9px;
  letter-spacing: 2px;
}
.publication-row .eyebrow {
  margin-bottom: 12px;
  font-size: 10px;
}
.publication-row h3 {
  font-size: 26px;
  line-height: 1.25;
  letter-spacing: -0.025em;
  margin-bottom: 14px;
}
.authors {
  font-size: 14px;
  color: var(--muted);
  line-height: 1.65;
}
.publication-summary {
  font-size: 15px;
  color: var(--muted);
}
.publication-links {
  display: flex;
  flex-wrap: wrap;
  gap: 8px 18px;
  font-size: 11px;
  margin-top: 18px;
}
.publication-links a {
  border-bottom: 1px solid var(--line);
  padding: 2px 0;
}
.home-news {
  padding-bottom: 100px;
}
.news-grid {
  display: grid;
  grid-template-columns: repeat(3, 1fr);
  gap: 40px;
}
.news-grid article {
  border-top: 1px solid var(--line);
  padding-top: 25px;
}
.news-grid h3 {
  font-size: 24px;
  margin-bottom: 15px;
}
.news-grid p:not(.eyebrow) {
  color: var(--muted);
  font-size: 14px;
}
.join-banner {
  padding: 58px 0 90px;
  border-top: 1px solid var(--line);
}
.join-banner h2 {
  font-size: clamp(48px, 6.5vw, 90px);
  margin-bottom: 28px;
  color: var(--teal);
}
.page-shell {
  padding-top: 80px;
  padding-bottom: 105px;
  min-height: 70vh;
}
.page-heading {
  margin-bottom: 65px;
}
.page-heading h1 {
  font-size: clamp(52px, 7vw, 96px);
  letter-spacing: -0.055em;
  margin-bottom: 25px;
}
.page-summary {
  font-size: 23px;
  line-height: 1.5;
  max-width: 800px;
  color: var(--muted);
  margin-bottom: 35px;
}
.methods-section {
  display: grid;
  grid-template-columns: 1fr 1fr;
  gap: 60px;
  padding: 70px 0 0;
}
.methods-section h2 {
  font-size: 32px;
}
.methods-section p {
  color: var(--muted);
}
.members-grid {
  display: grid;
  grid-template-columns: repeat(4, 1fr);
  gap: 55px 28px;
}
.member-card img,
.member-placeholder {
  aspect-ratio: 4/5;
  width: 100%;
  object-fit: cover;
  object-position: 50% 20%;
  background: var(--placeholder);
}
.member-card h3 {
  font-size: 23px;
  margin-top: 20px;
  margin-bottom: 7px;
}
.member-card p {
  font-size: 13px;
  color: var(--muted);
  line-height: 1.5;
}
.member-placeholder {
  display: flex;
  align-items: center;
  justify-content: center;
  font-size: 80px;
  letter-spacing: -5px;
  font-weight: 400;
  color: var(--placeholder-ink);
  background: linear-gradient(145deg, var(--placeholder), var(--tint));
}
.member-role {
  color: var(--teal) !important;
}
.alumni-section {
  margin-top: 90px;
  padding-top: 45px;
  border-top: 1px solid var(--line);
}
.alumni-section > p {
  color: var(--muted);
  margin: 20px 0 40px;
}
.alumni-grid {
  display: grid;
  grid-template-columns: repeat(3, 1fr);
  gap: 35px;
}
.alumni-grid h3 {
  font-size: 20px;
  margin-bottom: 8px;
}
.alumni-grid p {
  font-size: 13px;
  color: var(--muted);
}
.article-shell {
  padding: 60px 0 110px;
  min-height: 70vh;
}
.back-link {
  display: inline-block;
  color: var(--muted);
  font-size: 13px;
  margin-bottom: 45px;
}
.article-title {
  font-size: clamp(37px, 4.7vw, 66px);
  max-width: 1050px;
  line-height: 1.12;
  margin-bottom: 30px;
}
.article-figure {
  max-width: 850px;
  margin: 48px 0;
  padding: 24px;
  background: var(--surface);
}
.article-figure img {
  display: block;
  margin: auto;
  max-height: 600px;
  object-fit: contain;
}
.article-figure figcaption {
  font-size: 13px;
  color: var(--muted);
  padding-top: 20px;
}
.prose {
  font-size: 18px;
  line-height: 1.75;
  max-width: 800px;
  margin-top: 35px;
}
.prose h2 {
  font-size: 32px;
  margin: 40px 0 20px;
}
.prose h3 {
  font-size: 24px;
  margin: 25px 0 12px;
}
.prose a {
  color: var(--teal);
  border-bottom: 1px solid var(--line);
}
.prose img {
  display: block;
  margin: 30px auto;
  max-height: 600px;
  object-fit: contain;
}
.prose figure {
  margin: 40px 0;
}
.prose figcaption {
  font-size: 13px;
  color: var(--muted);
}
.related-publications {
  margin-top: 75px;
}
.related-publications > h2 {
  margin-bottom: 30px;
}
.profile-grid {
  display: grid;
  grid-template-columns: 300px 1fr;
  gap: 70px;
}
.profile-grid h1 {
  font-size: clamp(40px, 5vw, 68px);
  margin-bottom: 18px;
}
.profile-photo {
  width: 100%;
  aspect-ratio: 4/5;
  object-fit: cover;
  object-position: 50% 15%;
}
.profile-links {
  display: flex;
  gap: 20px;
  flex-wrap: wrap;
  font-size: 13px;
  color: var(--teal);
}
.publication-filters {
  display: grid;
  grid-template-columns: 2fr 1fr 1.5fr auto auto;
  align-items: end;
  gap: 22px;
  padding: 24px 0;
  border-top: 1px solid var(--line);
  border-bottom: 1px solid var(--line);
}
.publication-filters label {
  display: block;
  font-size: 11px;
  color: var(--muted);
  margin-bottom: 7px;
}
.publication-filters input[type="search"],
.publication-filters select {
  width: 100%;
  border: 0;
  border-bottom: 1px solid var(--muted);
  background: transparent;
  padding: 8px 0;
  font-size: 14px;
  border-radius: 0;
}
.publication-filters .featured-filter {
  white-space: nowrap;
  display: flex;
  align-items: center;
  gap: 8px;
  margin-bottom: 7px;
  font-size: 12px;
}
.featured-filter input {
  accent-color: var(--teal);
  width: 16px;
  height: 16px;
}
.reset-filters {
  font-size: 12px;
  background: none;
  border: 0;
  border-bottom: 1px solid var(--line);
  padding: 8px 0;
  margin-bottom: 1px;
}
.result-count {
  font-size: 12px;
  color: var(--muted);
  margin: 22px 0;
}
.news-row {
  display: grid;
  grid-template-columns: 150px 1fr 25px;
  gap: 30px;
  border-top: 1px solid var(--line);
  padding: 35px 0;
}
.news-row time {
  font-size: 13px;
  color: var(--muted);
}
.news-row h2 {
  font-size: 28px;
  margin-bottom: 18px;
}
.news-row p {
  color: var(--muted);
  font-size: 15px;
}
.news-image {
  max-height: 550px;
  object-fit: contain;
}
.join-layout {
  display: grid;
  grid-template-columns: 1.6fr 1fr;
  gap: 80px;
}
.join-layout .prose {
  margin-top: 0;
}
.join-layout .prose h2:first-child {
  margin-top: 0;
}
.contact-aside {
  align-self: start;
  background: var(--tint);
  padding: 38px 32px;
}
.contact-aside h2 {
  font-size: 35px;
  margin-bottom: 28px;
}
.contact-aside a {
  font-size: 14px;
  overflow-wrap: anywhere;
  color: var(--teal);
}
.contact-aside .small {
  margin-top: 30px;
  color: var(--muted);
}
.contact-grid {
  display: grid;
  grid-template-columns: minmax(0, 1.35fr) minmax(0, 1fr);
  gap: clamp(32px, 4vw, 64px);
  align-items: start;
}
.contact-building img {
  display: block;
  width: 100%;
  height: auto;
}
.contact-building figcaption {
  margin-top: 16px;
  color: var(--muted);
  font-size: var(--text-caption);
  line-height: 1.5;
}
.contact-details {
  min-width: 0;
}
.contact-details section + section {
  margin-top: 36px;
}
.contact-grid h2 {
  font-size: 30px;
  margin-bottom: 24px;
}
.contact-grid a {
  color: var(--teal);
}
.contact-grid p {
  margin: 20px 0;
  line-height: 1.6;
}
.contact-email {
  overflow-wrap: anywhere;
}
@media (max-width: 900px) {
  .contact-grid {
    grid-template-columns: 1fr;
    gap: 36px;
  }
}
@media (min-width: 1500px) {
  .story-stage {
    min-height: 740px;
  }
  .molecule-view {
    max-height: 720px;
  }
}
@media (max-width: 1050px) {
  .wrap {
    width: calc(100% - 64px);
  }
  .site-header {
    padding-inline: 32px;
  }
  .site-header nav {
    gap: 18px;
  }

  .hero-copy h1 {
    font-size: 9.1vw;
  }
  .hero-intro {
    max-width: 290px;
    font-size: 16px;
  }
  .molecule-label {
    font-size: 10px;
  }
  .label-receptor {
    right: 0;
  }
  .story-step {
    max-width: 33vw;
  }
  .publication-row {
    grid-template-columns: 200px 1fr;
    gap: 30px;
  }
  .members-grid {
    grid-template-columns: repeat(3, 1fr);
  }
  .publication-filters {
    grid-template-columns: 2fr 1fr 1.5fr;
    gap: 20px;
  }
  .featured-filter,
  .reset-filters {
    justify-self: start;
  }
  .mission-section {
    grid-template-columns: 1fr 2fr;
    gap: 30px;
  }
  .join-layout {
    gap: 40px;
  }
  .research-row {
    grid-template-columns: 1fr 1fr 20px;
    gap: 20px;
  }
}
@media (max-width: 700px) {
  body {
    font-size: 16px;
  }
  .wrap {
    width: calc(100% - 40px);
  }
  .site-header {
    padding: 22px 20px;
    min-height: 79px;
    flex-wrap: wrap;
  }
  .wordmark {
    font-size: clamp(19px, 5vw, 23px);
    gap: 6px;
  }
  .wordmark span {
    display: inline;
    font-size: inherit;
  }
  .site-header nav {
    width: 100%;
    flex-wrap: wrap;
    gap: 8px 22px;
    padding-top: 20px;
  }
  .js .menu-toggle {
    display: block;
  }
  .js .site-header nav:not(.is-open) {
    display: none;
  }
  .site-header nav a {
    font-size: 14px;
  }
  .hero-copy {
    padding: 34px 0 0;
  }
  .hero-copy .eyebrow {
    font-size: 9px;
    margin-bottom: 21px;
  }
  .hero-copy h1 {
    font-size: clamp(62px, 15vw, 98px);
    line-height: 0.97;
  }
  .hero-intro {
    max-width: 290px;
    font-size: 15px;
    margin: 22px 0;
  }
  .scroll-cue {
    font-size: 12px;
  }
  .story-stage {
    display: block;
    min-height: 0;
  }
  .molecular-figure {
    margin: 18px 0 35px;
  }
  .molecule-view {
    max-width: 500px;
    max-height: 430px;
    margin: auto;
  }
  .molecule-label {
    font-size: 10px;
  }
  .figure-note {
    font-size: 9px;
  }
  .molecular-figure figcaption {
    font-size: 9px;
  }
  .motion-enabled .story-stage {
    position: relative;
    height: auto;
    min-height: 0;
  }
  .motion-enabled .story-stage .molecular-figure {
    position: sticky;
    top: 10px;
  }
  .motion-enabled .story-steps {
    margin-top: 0;
  }
  .motion-enabled .story-step {
    min-height: 360px;
    max-width: 100%;
    padding: 40px 0;
  }
  .motion-enabled .molecular-story {
    padding-bottom: 0;
  }
  .motion-enabled .hero-copy {
    opacity: 1 !important;
  }
  .story-step {
    max-width: 100%;
    padding: 40px 0;
  }
  .story-step h2 {
    font-size: 40px;
  }
  .story-step p {
    max-width: 500px;
  }
  .mission-section {
    padding: 55px 0;
    display: block;
  }
  .mission-section h2 {
    font-size: 33px;
  }
  .mission-section .large-copy {
    font-size: 20px;
  }
  .section-heading h2 {
    font-size: 30px;
  }
  .section-heading .text-link {
    font-size: 12px;
    gap: 10px;
  }
  .section-heading {
    gap: 20px;
    align-items: center;
  }
  .research-preview,
  .selected-publications,
  .home-news {
    padding-bottom: 60px;
  }
  .research-row {
    grid-template-columns: 1fr 20px;
    gap: 15px;
    padding: 27px 0;
  }
  .research-row h2,
  .research-row h3 {
    font-size: 25px;
  }
  .research-row p {
    grid-column: 1;
    grid-row: 2;
  }
  .research-row > span:last-child {
    grid-column: 2;
    grid-row: 1;
  }
  .publication-row {
    grid-template-columns: 1fr;
    gap: 25px;
    padding: 28px 0;
  }
  .publication-image {
    max-height: 230px;
    min-height: 0;
    padding: 18px;
  }
  .publication-image img {
    max-height: 180px;
  }
  .publication-year-art {
    display: none;
  }
  .publication-row h3 {
    font-size: 24px;
  }
  .news-grid {
    grid-template-columns: 1fr;
    gap: 25px;
  }
  .news-grid h3 {
    font-size: 25px;
  }
  .join-banner {
    padding: 45px 0 60px;
  }
  .join-banner h2 {
    font-size: 49px;
  }
  .site-footer {
    padding: 35px 0;
  }
  .footer-brand {
    margin-bottom: 32px;
  }
  .footer-grid {
    grid-template-columns: 1fr;
    gap: 25px;
  }
  .footer-grid > div:last-child {
    text-align: left;
  }
  .site-footer p {
    margin: 5px 0;
  }
  .page-shell {
    padding: 48px 0 70px;
  }
  .page-heading {
    margin-bottom: 40px;
  }
  .page-heading h1 {
    font-size: 56px;
  }
  .page-summary {
    font-size: 20px;
  }
  .methods-section {
    display: block;
    padding-top: 45px;
  }
  .methods-section h2 {
    margin-bottom: 20px;
  }
  .members-grid {
    grid-template-columns: repeat(2, 1fr);
    gap: 36px 18px;
  }
  .member-card h3 {
    font-size: 21px;
  }
  .member-placeholder {
    font-size: 55px;
  }
  .alumni-grid {
    grid-template-columns: 1fr 1fr;
    gap: 25px;
  }
  .alumni-section {
    margin-top: 60px;
  }
  .article-shell {
    padding: 35px 0 70px;
  }
  .article-title {
    font-size: 37px;
  }
  .back-link {
    margin-bottom: 30px;
  }
  .profile-grid {
    grid-template-columns: 1fr;
    gap: 35px;
  }
  .profile-photo,
  .profile-placeholder {
    max-width: 280px;
  }
  .profile-grid h1 {
    font-size: 45px;
  }
  .prose {
    font-size: 17px;
  }
  .publication-filters {
    grid-template-columns: 1fr 1fr;
    gap: 18px;
  }
  .search-field {
    grid-column: 1/-1;
  }
  .publication-filters select {
    font-size: 13px;
  }
  .publication-filters .featured-filter {
    grid-column: 1;
  }
  .news-row {
    grid-template-columns: 1fr 20px;
    gap: 15px;
  }
  .news-row time {
    grid-column: 1/-1;
  }
  .news-row h2 {
    font-size: 26px;
  }
  .join-layout,
  .contact-grid {
    grid-template-columns: 1fr;
    gap: 40px;
  }
  .contact-aside {
    padding: 30px;
  }
  .contact-grid h2 {
    font-size: 27px;
  }
}
@media (prefers-reduced-motion: reduce) {
  html {
    scroll-behavior: auto;
  }
  *,
  *:before,
  *:after {
    transition: none !important;
    animation: none !important;
  }
}
@media print {
  .site-header,
  .site-footer,
  .publication-filters,
  .motion-toggle {
    display: none;
  }
  .wrap {
    width: 100%;
  }
  .molecular-story {
    display: none;
  }
  a {
    color: inherit;
  }
  .publication-row {
    break-inside: avoid;
  }
  .page-shell {
    padding: 0;
  }
}
/* On small screens the molecular figure stays above the scrolling story. */
@media (max-width: 700px) {
  .motion-enabled .story-stage {
    display: contents;
  }
  .motion-enabled .hero-copy {
    width: calc(100% - 40px);
    margin-inline: auto;
  }
  .motion-enabled .story-stage > .molecular-figure {
    position: sticky;
    top: 0;
    z-index: 6;
    width: calc(100% - 40px);
    margin: 20px auto 0;
    padding: 8px 0 6px;
    background: var(--paper);
  }
  .motion-enabled .molecule-view {
    height: 39svh;
    max-height: 350px;
    width: 100%;
    aspect-ratio: auto;
  }
  .motion-enabled .molecular-figure figcaption {
    padding: 7px 0;
    gap: 3px;
  }
  .motion-enabled .figure-note {
    display: none;
  }
  .motion-enabled .motion-toggle {
    bottom: 0;
    right: 0;
    transform: translateY(100%);
    background: var(--paper);
    padding: 3px 5px;
  }
  .motion-enabled .story-step {
    min-height: 65svh;
    padding: 60px 0 40px;
  }
  .motion-enabled .story-step h2 {
    font-size: 36px;
  }
  .motion-enabled .story-step .eyebrow {
    margin-bottom: 14px;
  }
}

/* Reading scale: the mission paragraph is the reference for all regular text.
   Keep these tokens together so future typography changes are straightforward. */
:root {
  --text-body: 23px;
  --text-secondary: 20px;
  --text-ui: 17px;
  --text-caption: 16px;
  --text-label: 14px;
}
body,
.hero-intro,
.story-step p,
.mission-section .large-copy,
.research-row p,
.publication-summary,
.news-grid p:not(.eyebrow),
.page-summary,
.methods-section p,
.member-card p,
.alumni-grid p,
.prose,
.news-row p {
  font-size: var(--text-body);
  line-height: 1.5;
}
.hero-intro {
  max-width: 410px;
}
.prose {
  line-height: 1.6;
}
.authors {
  font-size: var(--text-secondary);
}
small,
.small,
.article-figure figcaption,
.prose figcaption,
.site-footer,
.publication-links,
.profile-links,
.back-link,
.result-count,
.news-row time {
  font-size: var(--text-caption);
}
.site-header nav,
.site-header nav a,
.menu-toggle,
.text-link,
.section-heading .text-link,
.publication-filters input[type="search"],
.publication-filters select,
.publication-filters .featured-filter,
.reset-filters {
  font-size: var(--text-ui);
}
.publication-filters label {
  font-size: var(--text-caption);
}
.eyebrow,
.hero-copy .eyebrow,
.publication-row .eyebrow,
.publication-year-art span {
  font-size: var(--text-label);
}
.molecule-label {
  font-size: var(--text-caption);
}
.molecule-label small,
.molecular-figure figcaption,
.figure-index,
.figure-note,
.motion-toggle {
  font-size: var(--text-label);
}
.contact-aside a,
.footer-name {
  font-size: var(--text-secondary);
}
.research-row h2,
.research-row h3 {
  font-size: 32px;
}
.member-card h3,
.alumni-grid h3,
.news-grid h3 {
  font-size: 27px;
}
.publication-row h3 {
  font-size: 29px;
}
.menu-toggle {
  display: none;
}
/* Do not compress long titles or controls when copy wraps. */
.research-row > *,
.publication-row > *,
.members-grid > *,
.alumni-grid > *,
.news-grid > *,
.publication-filters > * {
  min-width: 0;
}
.publication-links {
  gap: 10px 20px;
}
@media (max-width: 1050px) {
  .site-header nav {
    gap: 16px;
  }
}
@media (max-width: 700px) {
  :root {
    --text-body: 20px;
    --text-secondary: 18px;
  }
  .hero-intro {
    max-width: 100%;
  }
  .research-row h2,
  .research-row h3 {
    font-size: 28px;
  }
  .member-card h3,
  .alumni-grid h3,
  .news-grid h3 {
    font-size: 25px;
  }
  .publication-row h3 {
    font-size: 27px;
  }
  .section-heading {
    flex-wrap: wrap;
    gap: 16px;
  }
  .publication-filters .featured-filter {
    white-space: normal;
  }
  .molecule-label {
    font-size: 14px;
  }
  .molecule-label small {
    font-size: 12px;
  }
}
@media (max-width: 420px) {
  .members-grid,
  .alumni-grid {
    grid-template-columns: 1fr;
  }
  .member-card img,
  .member-placeholder {
    max-width: 300px;
  }
}

/* Visitor display preferences; defaults live in _data/display.yml. */
.display-settings {
  position: relative;
  padding-block: 0;
  font-size: var(--text-ui);
}
.display-settings summary {
  cursor: pointer;
  display: grid;
  place-items: center;
  list-style: none;
  width: 44px;
  height: 44px;
  color: var(--ink);
  white-space: nowrap;
}
.display-settings summary::-webkit-details-marker {
  display: none;
}
.display-settings[open] .settings-chevron {
  transform: rotate(180deg);
}
.display-settings summary:hover {
  background: var(--tint);
  border-radius: 5px;
}
.molstar-controls {
  margin: 0;
  padding: 0;
  border: 0;
}
.molstar-controls legend {
  margin-bottom: 10px;
}
.display-controls .display-check {
  display: flex;
  align-items: center;
  gap: 10px;
  margin-block: 8px;
}
.display-check input {
  width: 18px;
  height: 18px;
  margin: 0;
  accent-color: var(--teal);
}
.ssgi-support-note {
  color: var(--muted);
  font-size: 14px;
}
.display-settings summary:focus-visible {
  outline: 2px solid var(--teal);
  outline-offset: 4px;
}
.display-controls {
  position: absolute;
  top: calc(100% + 12px);
  right: 0;
  width: 350px;
  max-width: calc(100vw - 40px);
  display: grid;
  gap: 18px;
  padding: 22px;
  border: 1px solid var(--line);
  border-radius: 8px;
  background: var(--paper);
  box-shadow: 0 12px 35px rgb(0 0 0 / 12%);
}
.display-reset {
  justify-self: start;
}
@media (min-width: 701px) and (max-width: 1180px) {
  .site-header {
    flex-wrap: wrap;
  }
  .js .menu-toggle {
    display: block;
  }
  .site-header nav {
    width: 100%;
    padding-top: 20px;
    flex-wrap: wrap;
  }
  .js .site-header nav:not(.is-open) {
    display: none;
  }
}
@media (max-width: 1180px) {
  .display-settings {
    width: 100%;
  }
  .display-controls {
    position: static;
    margin-top: 12px;
    box-shadow: none;
    width: min(100%, 420px);
  }
}

.display-controls label {
  min-width: 0;
  display: grid;
  gap: 5px;
}
.display-controls select {
  width: 100%;
  min-width: 0;
  max-width: 100%;
  border: 1px solid var(--line);
  border-radius: 4px;
  background: var(--paper);
  padding: 8px 30px 8px 10px;
}
.display-reset {
  background: none;
  border: 0;
  border-bottom: 1px solid var(--line);
  padding: 8px 0;
}
.molecule-renderer {
  position: absolute;
  inset: 0;
  pointer-events: none;
}
.molecule-renderer canvas,
.pymol-frame,
.pymol-dynamics-layer {
  position: absolute;
  inset: 0;
  width: 100%;
  height: 100%;
  object-fit: contain;
}
.molecule-renderer > div {
  position: absolute;
  inset: 0;
}
.pymol-frame {
  opacity: 0;
}
.molecule-ready .pymol-frame {
  opacity: 1;
}
.renderer-status {
  font-size: var(--text-label);
  color: var(--muted);
  margin: 0;
  min-height: 0;
}
.renderer-status:empty {
  display: none;
}
.label-peptide:before {
  background: var(--peptide_color);
}
.label-ligand:before {
  background: var(--ligand_color);
}
.label-receptor:before {
  background: var(--receptor_color);
}
@media (max-width: 700px) {
  .display-controls {
    display: grid;
    grid-template-columns: 1fr;
  }
  .display-reset {
    justify-self: start;
  }
}

/* Reserve room for the caption and pause control on short desktop windows. */
@media (min-width: 701px) {
  .motion-enabled .molecule-view {
    max-height: calc(100svh - 160px);
  }
}